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The Medial Hemispheres Are Involved in Adjusting Limb Movements The major inputs to the medial hemispheres (paravermal cortex) are superimposed, somatotopically arranged projections from the motor cortex and spinal cord (see Fig. 20-16 ). The major output of this part of the cerebellum is via the interposed nucleus back to motor cortex (through VL/VA; see Fig. 20-22B ) and to the magnocellular red nucleus. Thus the medial hemispheres can influence spinal cord motor neurons through the corticospinal tract and also, to a minor extent, through the rubrospinal pathway. This led to the hypothesis that the intermediate zone of the cerebellum compares the commands emanating from motor cortex (it receives this information via pontine nuclei) with the actual position and velocity of the moving part (it receives this information via spinocerebellar and similar tracts) and then, by way of the interposed nucleus, issues correcting signals. This is consistent with the observation that most neurons of the interposed nucleus have firing rates related to voluntary movements, but unlike those of dentate neurons, their rates tend to change during rather than before movement ( Fig. 20-23 ). Note that here again, a given side of the cerebellum winds up affecting ipsilateral motor neurons (e.g., left side of cerebellum→right thalamus→right motor cortex→left side of spinal cord).
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* השאלה נוספה בתאריך: 12-07-2020